The Crisis On Infinite Earths was the ridiculous solution that DC Comics found to explain the contradicting and confusing continuity that happens when you write stories in a conjoined universe for forty years. The conflict began with the Anti-Monitor, a nearly omnipotent being created when an Oan scientist tried look back in time to watch the Big Bang and accidentally created the DC multiverse.
Don't go anywhere, we're just skimming the surface of continuity crack here.
The Anti-Monitor wants to destroy every universe in the multiverse, and has so far been successful: only five remain. His counterpart, the Monitor, recruits heroes to defend a bunch of giant tuning forks and keep the Anti-Monitor from messing everything up long enough to snap all the surviving parallel Earths into one Earth, which will be able to survive the Anti-Monitor's assault.
I tried to come up with a good summary for the rest of this, but lets just say: And then a bunch of shit happened. (Like the epic deaths of the first Supergirl and Flash.)
Eventually, the Spectre and the Anti-Monitor wind up going mano-a-mano and the force of their battle shatters space-time, creating a single universe where all five DC Earths are merged and no one remembers that anything was ever any different.
tl;dr: The writers rewrote the rules so that they could do whatever they wanted with DC characters from now on. Until Grant Morrison.
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